Member list
Current members
- Lan-Wei Yeh
- Ecology and evolution of Taiwanese Carabus ground beetles.
- Chung-Hsin Huang
- Master topic: Flucuating Asymmetry and Developmental Asymmetry of Cyclommatus mniszechi; PhD topic: Evolution of Mandible Morphology and Fighting Behavior in Stag Beetles
- Ying-Yuan Lo
- Systematics, Behavior and Evolution of Lynx Spiders.
- Leocris Batucan Jr.
- New Species and DNA Barcoding of the Mayfly (Ephemeroptera) in Mt. Malindang, Mindanao, Philippines.
- Chun-Yu Lin
- Parasitoid Wasps of Damselflies; Phylogenetics and Evolution Histories of Treehoppers
- Hsu-Yi Lin
- Fighting Behavior of Odontolabis siva parryi Stag Beetles
- Sheng Lin
- Transcriptome Profiling Psolodesmus mandarinus mandarinus Damselfly
- Pei-Hsuan Chang
- Thermal Adaptation and Habitat Preference of Psolodesmus mandarinus Damselfly
- Min-Chen Lin
- Disruptive Camauflage of Colors of Pachyrhynchus Weevils.
- Masahiko Tanahashi
- Ecology and Evolution of Pachyrhynchus Weevils.
- Su-Ping Lin
- Symbiotic Yeasts of Stag Beetles
- Jun-Ying Wang
- Conservation of Mortonagrion hirosei Damselflies
- Chin-Han Hsiao
- Fighting Behavior of Rhaetulus crenatus Stag Beetles
- Wei-Zhe Tseng
- Systematics and Evolution of Weevils
- Cheng-Tse Yung
- Maternal Gift of Pachyrhynchus sarcitis Weevils
Past members
- Jo-Fan Wang
- Territorial behavior, trait evolution.
- Yu-Hsun Hsu
- Mating strategy, behavioural ecology, molecular ecology, population genetics, molecular phylogenetics, biostatistics.
- Yi-Shou Yang
- Reproductive Isolation of Psolodesmus mandarinus subspecies.
- Wataru Kojima
- Zhen-Yi Chen
- Fighting behavior and reproductive strategy of stag beetles.
- Teng-Yu Liu
- Fighting behaviour and life history observation of a stag beetle Rhaetulus crenatus crenatus.
- Princess Angelie S. Casas
- Insect Diversity and Evolution
- Lu-Yi Wang
- Defense Mechanism of Pachyrhynchus weevils.
- Hui-Yun Tseng
- Aposematic coloration, island biogeography.
- Yu-Jen Yu
- Variation and Adaptation of wings.
- Yen-Ting Chen
- Systematics and evolution of insects.
- Yun-Chieh Cheng
- Damselfly larval diet, molecular evolution.
- Tai-Chia Chiang
- The effect of aposematic colors of Pachyrhynchus weevils on avian predators.
- Yong-Chao Su
- Behavioral ecology, sociobiology, population genetics, and molecular phylogenetics.
- I-Ting Hsiao
- Variation of genitalia in Euphaea amphicyana.
- Jyun-Huei Huang
- The fighting behavior of a stag beetle, Rhaetulus crenatus.
- Shi-Ting Wu
- Membracis Phyllotropis.
- Chiao-Wei Lin
- The fighting behavior of a stag beetle, Rhaetulus crenatus.
- Wei-Liang Xiao
- Variation of wing veins in Euphaea amphicyana.
- Hung-Nien Chen
- 1. The effect of typhoon on survivalship of Matrona cyanoptera. 2. Modes of phenotypic variation in Euphaea amphicyana.
- Chu-Yen Cheng
- Phylogeography of a Philippine's treehopper, Leptocentrus reponens.
- Che-Yu Kuan
- Variation of mandibles in stag beetles.
- Yat-Hung Lee
- Speciation of Euphaea damselflies.
- Ming-Yu Chen
- Phylogeography and population history of the treehoppers, Centrochares horifficus from the Philippines Archipelago.
- Shao-Chang Huang
- Visual Communication of Matrona cyanoptera
- Jen-Pan Huang
- Population Genetics and Phylogeographic Analyses of Formosan Damselfly, Euphaea Formosa (Insecta: Odonata: Euphaeidae) from Taiwan
- Wei-Yun Chen
- Molecular and Phylogenetic Characterization of Endosymbiotic Bacteria of the Froghopper, Okiscarta uchidae (Insecta: Hemiptera: Cercopidae)
Visiting scholars and students
- Jillian del Sol
- Behavioral Ecology; Mating System Ecology; Weapon Ecology and Evolution; Evolutionary Biology
- Ian Dela Cruz
- New Species and DNA Barcoding of the Stonefly (Plecoptera) in Mt. Malindang, Mindanao, Philippines.
- Shinya Komata
- Ecology & Evolution of Wing Polymorphism in a Mimetic Swallowtail Butterfly, Papilio memnon.
- Vanitha Williams
- Predatory potential of waterbug, Diplonychus rusticus and dragonfly, Diplacodes trivialis on mosquito larvae.
- Marina Vilenica
- Dragonfly composition (Insecta, Odonata) in wetland area of Turopolje region, Croatia
- Klaas-Douwe 'KD' B. Dijkstra
- History, diversity and identification of dragonflies and damselflies (Odonata).
- Erin McCullough
- " Diversification of weapon form: aerodynamic costs of beetle horns.
- Ashley E. King
- Intrasexual combat and intersexual antagonistic co-evolution in horned beetles.
Wataru Kojima

Title
JSPS Postdoctoral Fellow for Research Abroad 2016-
Species: Dicronocephalus wallichii, Trypoxylus dichotomus, Cheirotonus formosanus

Project
Evolution of male sexual traits in Taiwanese scarab beetles.
Abstrct
E-mail:koj.wataru@gmail.com
Males of some Taiwanese scarab beetles have exaggerated traits including horns and enlarged forelegs. These traits may have evolved through sexual selection, but the function of them is largely unknown. I will examine their function in mate guarding and intrasexual competition. I will also examine geographic variations in these traits and identify the ecological factors responsible for the variations.
The males of a flower beetle Dicronocephalus bourgoini are equipped with exaggerated horns and elongated forelegs. We investigated their functions by examining male-male competition in the field. Our observation showed that the sex ratio of this species at feeding/mating sites was male-biased and the competition between the mate-guarding (i.e., owners) and unpaired males (i.e., intruders) frequently occurred. In the early phase of the contests, males mainly used their forelegs, likely to assess the body size of their opponents. If the foreleg length of the owner was shorter than that of the intruder, the intruder approached to take over the owner’s mate, causing escalated contests to occur, in which both opponents tried to drag each other away from the females or the substrate using their horns. About a half of all contests were terminated before escalating to direct combat. These results suggest that the multiple weapons in this species are specialized for specific phases of contests.

A mate-guarding male (upper) extended his forelegs and tapped an intruder (lower) of Dicronocephalus bourgoini.