Member list
Current members
- Lan-Wei Yeh
- Ecology and evolution of Taiwanese Carabus ground beetles.
- Chung-Hsin Huang
- Master topic: Flucuating Asymmetry and Developmental Asymmetry of Cyclommatus mniszechi; PhD topic: Evolution of Mandible Morphology and Fighting Behavior in Stag Beetles
- Ying-Yuan Lo
- Systematics, Behavior and Evolution of Lynx Spiders.
- Leocris Batucan Jr.
- Chun-Yu Lin
- Parasitoid Wasps of Damselflies; Phylogenetics and Evolution Histories of Treehoppers
- Hsu-Yi Lin
- Fighting Behavior of Odontolabis siva parryi Stag Beetles
- Sheng Lin
- Transcriptome Profiling Psolodesmus mandarinus mandarinus Damselfly
- Pei-Hsuan Chang
- Thermal Adaptation and Habitat Preference of Psolodesmus mandarinus Damselfly
- Min-Chen Lin
- Disruptive Camauflage of Colors of Pachyrhynchus Weevils.
- Masahiko Tanahashi
- Ecology and Evolution of Pachyrhynchus Weevils.
- Su-Ping Lin
- Symbiotic Yeasts of Stag Beetles
- Jun-Ying Wang
- Conservation of Mortonagrion hirosei Damselflies
- Chin-Han Hsiao
- Fighting Behavior of Rhaetulus crenatus Stag Beetles
- Wei-Zhe Tseng
- Systematics and Evolution of Weevils
- Cheng-Tse Yung
- Maternal Gift of Pachyrhynchus sarcitis Weevils
Past members
- Jo-Fan Wang
- Territorial behavior, trait evolution.
- Yu-Hsun Hsu
- Mating strategy, behavioural ecology, molecular ecology, population genetics, molecular phylogenetics, biostatistics.
- Yi-Shou Yang
- Reproductive Isolation of Psolodesmus mandarinus subspecies.
- Wataru Kojima
- Evolution of male sexual traits in Taiwanese scarab beetles.
- Zhen-Yi Chen
- Fighting behavior and reproductive strategy of stag beetles.
- Teng-Yu Liu
- Fighting behaviour and life history observation of a stag beetle Rhaetulus crenatus crenatus.
- Princess Angelie S. Casas
- Insect Diversity and Evolution
- Lu-Yi Wang
- Defense Mechanism of Pachyrhynchus weevils.
- Hui-Yun Tseng
- Aposematic coloration, island biogeography.
- Yu-Jen Yu
- Variation and Adaptation of wings.
- Yen-Ting Chen
- Systematics and evolution of insects.
- Yun-Chieh Cheng
- Damselfly larval diet, molecular evolution.
- Tai-Chia Chiang
- The effect of aposematic colors of Pachyrhynchus weevils on avian predators.
- Yong-Chao Su
- Behavioral ecology, sociobiology, population genetics, and molecular phylogenetics.
- I-Ting Hsiao
- Variation of genitalia in Euphaea amphicyana.
- Jyun-Huei Huang
- The fighting behavior of a stag beetle, Rhaetulus crenatus.
- Shi-Ting Wu
- Membracis Phyllotropis.
- Chiao-Wei Lin
- The fighting behavior of a stag beetle, Rhaetulus crenatus.
- Wei-Liang Xiao
- Variation of wing veins in Euphaea amphicyana.
- Hung-Nien Chen
- 1. The effect of typhoon on survivalship of Matrona cyanoptera. 2. Modes of phenotypic variation in Euphaea amphicyana.
- Chu-Yen Cheng
- Phylogeography of a Philippine's treehopper, Leptocentrus reponens.
- Che-Yu Kuan
- Variation of mandibles in stag beetles.
- Yat-Hung Lee
- Speciation of Euphaea damselflies.
- Ming-Yu Chen
- Phylogeography and population history of the treehoppers, Centrochares horifficus from the Philippines Archipelago.
- Shao-Chang Huang
- Visual Communication of Matrona cyanoptera
- Jen-Pan Huang
- Population Genetics and Phylogeographic Analyses of Formosan Damselfly, Euphaea Formosa (Insecta: Odonata: Euphaeidae) from Taiwan
- Wei-Yun Chen
- Molecular and Phylogenetic Characterization of Endosymbiotic Bacteria of the Froghopper, Okiscarta uchidae (Insecta: Hemiptera: Cercopidae)
Visiting scholars and students
- Jillian del Sol
- Behavioral Ecology; Mating System Ecology; Weapon Ecology and Evolution; Evolutionary Biology
- Ian Dela Cruz
- New Species and DNA Barcoding of the Stonefly (Plecoptera) in Mt. Malindang, Mindanao, Philippines.
- Shinya Komata
- Ecology & Evolution of Wing Polymorphism in a Mimetic Swallowtail Butterfly, Papilio memnon.
- Vanitha Williams
- Predatory potential of waterbug, Diplonychus rusticus and dragonfly, Diplacodes trivialis on mosquito larvae.
- Marina Vilenica
- Dragonfly composition (Insecta, Odonata) in wetland area of Turopolje region, Croatia
- Klaas-Douwe 'KD' B. Dijkstra
- History, diversity and identification of dragonflies and damselflies (Odonata).
- Erin McCullough
- " Diversification of weapon form: aerodynamic costs of beetle horns.
- Ashley E. King
- Intrasexual combat and intersexual antagonistic co-evolution in horned beetles.
Leocris Batucan Jr.

Title
Master student, Mindanao State University-Iligan Institute of Technology, Mindanao, the Philippines (MECO-TECO exchange student 2014-2015); TIGP Biodiversity PhD Student 2016-
Species: Mayflies, Psolodesmus mandirinus Damselflies

Project
New Species and DNA Barcoding of the Mayfly (Ephemeroptera) in Mt. Malindang, Mindanao, Philippines.
Abstrct
A sucker for information, Leo likes to read write-ups about anything the world has to offer - thus daily giving him the sleepy/tired eyes.
E-mail:leocrisjr@gmail.com
Layawan River of Mt. Malindang in Mindanao Island

New Species and DNA Barcoding of the Mayfly (Ephemeroptera) in Mt. Malindang, Mindanao, Philippines
A major waterway in Mt. Malindang Range, Layawan River sustains many plants and animals. The project is aimed to determine the mayfly (Ephemeroptera) diversity of the river which is believed to have many un-described invertebrates. Geared to being a baseline study, the project described and identified several mayfly species and looks forward to DNA barcode of the specimens for biodiversity conservation efforts and other future works.
Novel wing display and divergent agonistic behaviors of two incipient Psolodesmus damselflies

Psolodesmus mandarinus dorothea (a) male and (b) female with small black apical pigmentation and hyaline wings, and Psolodesmus mandarinus mandarinus (c) male and (d) female with larger pigmented area, conspicuous white patch, and brown basal coloration.
Sexual selection via male competition is a strong evolutionary force that can drive rapid changes in competitive traits and subsequently lead to population divergence and speciation. Territorial males of many odonates are known to use their colorful wings as visual signals and to perform agonistic displays toward intruders. Psolodesmus mandarinus dorothea and Psolodesmus mandarinus mandarinus are two parapatrically distributed sister damselflies that share similar ecological characteristics but differ markedly in wing coloration. The wings of P. m. dorothea are mostly clear, whereas those of P. m. mandarinus have a large area of black pigmentation and a central white patch. We investigated whether territorial males of the two damselflies at breeding sites display distinct agonistic behaviors associated with their respective wing colors. Behavioral interactions between territorial and intruder males and their wing kinematics were filmed and analyzed for P. m. dorothea in Lienhuachih of central Taiwan, and P. m. mandarinus in Tianxiyuan and Fusan of northern Taiwan. We observed that the P. m. mandarinus males exhibited a novel set of perched wing displays, which was not only absent in its sister P. m. dorothea but also previously unknown in Odonata. At breeding sites, perched rival males of P. m. mandarinus with pigmented wings exhibited escalating agonistic wing-flapping and wing-hitting displays toward each other. In contrast, territorial males of P. m. dorothea with clear wings engaged only in aerial chase or face-to-face hovering when intruder males approached from the air. These results indicate that the two sister P. mandarinus damselflies diverged behaviorally in territorial contests and support the hypothesis of coadaptation on the basis of wing colors and types of wing movement in Odonata. Our findings further suggest that divergent agonistic wing displays may play a pivotal role in the speciation mechanism of P. mandarinus damselflies. The sequential analyses of behavioral characteristics and progression suggest that P. m. mandarinus damselflies likely use mutual assessment of rivals in territorial contests.

Ethograms of territorial contests of (a) Psolodesmus mandarinus dorothea and (b) Psolodesmus mandarinus mandarinus.